Set after the events of Continental Drift, the epic search for his acorn that is elusive of Scrat catapults him out of Earth, where he sets off a collection of events that transform and endanger the planet. To rescue themselves out of danger, Manny, Sid, Diego, and the remainder of the herd leave their house and embark on a quest full of thrills and spills, highs and lows, laughter and adventure when visiting exotic new lands and limiting a range of colorful new characters.